Trager Award



To honour his distinguished career in protozoology and his leadership role in establishing The Journal of Protozoology, the Society of Protozoologists established in 1998 the William Trager Award for Outstanding Paper of the Year. This award is given to the most cited original article published in the journal in the previous two years and consists of a Diploma and a $ 100.00 check to the first author of the article.

Recent past awardees:

Xavier Pochon and Todd C. LaJeunesse
Miliolidium n. gen, a New Symbiodiniacean Genus Whose Members Associate with Soritid Foraminifera or Are Free-Living” (vol. 68, e12856, 2021,

Edel Pérez-López, Md Musharaf Hossain, Jiangying Tu, Matthew Waldner, Christopher D. Todd, Anthony J. Kusalik, Yangdou Wei, Peta C. Bonham-Smith
Transcriptome Analysis Identifies Plasmodiophora brassicae Secondary Infection Effector Candidates” (vol. 67, pp. 337-351, 2020,

Allison M. Lewis, Andrea N. Chan, Todd C. LaJeunesse
New Species of Closely Related Endosymbiotic Dinoflagellates in the Greater Caribbean have Niches Corresponding to Host Coral Phylogeny” (vol 66, Issue 3, pp. 460-482,

David Bass, Lucas Czech, Bryony A. P. Williams, Cédric Berney, Micah Dunthorn, Frederic Mahé, Guifré Torruella, Grant D. Stentiford, Tom A. Williams
Clarifying the Relationships between Microsporidia and Cryptomycota”, (vol. 65, Issue 6, pp. 773-782,

Kensuto Seto, Maiko Kagami, and Yosuke Degawa
Phylogenetic Position of Parasitic Chytrids on Diatoms: Characterization of a Novel Clade in Chytridiomycota” (vol. 64, Issue 3, pp. 383-393,

Meng Qi, Rongjun Wang, Haiyan Wang, Fuchun Jian, Junqiang Li, Jinfeng Zhao, Haiju Dong, Huili Zhu, Changshen Ning and Longxian Zhang
Enterocytozoon bieneusi genotypes in grazing horses in China and their zoonotic transmission potential” (vol. 63, Issue 5, pp. 591-97).

Ryan S. Schwarz, Gary R. Bauchan, Charles A. Murphy, Jorgen Ravoet, Dirk C. de Graaf and Jay D. Evans
Characterization of two species of trypanosomatidae from the honey bee Apis mellifera: Crithidia mellificae Langridge and McGhee, 1967 and Lotmaria passim n. gen, n. sp.” (vol. 62, Issue 5, pp. 567-83).

Hae Jin Jeong, Sung Yeon Lee, Nam Seon Kang, Yeong Du Yoo, An Suk Lim, Moo Joon Lee, Hyung Seop Kim, Wonho Yih, Hiroshi Yamashita and Todd C. LaJeunesse
Genetics and Morphology Characterize the Dinoflagellate Symbiodinium voratum, n. sp., (Dinophyceae) as the Sole Representative of Symbiodinium Clade E” (vol. 61, Issue 1, pp. 75-94).

Yvonne Qvarnstrom, Thomas A. Nerad and Govinda S. Visvesvara
Characterization of a New Pathogenic Acanthamoeba Species, A. byersi n. sp., Isolated from a Human with a Fatal Amoebic Encephalitis” (vol. 60, Issue 6, pp. 626-633, 2013)

John R. Barta, Joseph D. Ogedengbe, Donald S. Martin and Todd G. Smith
Phylogenetic Position of the Adeleorinid Coccidia (Myzozoa, Apicomplexa, Coccidia, Eucoccidiorida, Adeleorina) Inferred Using 18S rDNA Sequences” (vol. 59, Issue 2, pp. 171-180)

Tsvetan R. Bachvaroff, Sara M., Handy, Allen R. Place and Charles F. Delwiche
Alveolate Phylogeny Inferred using Concatenated Ribosomal Proteins” (vol. 58, Issue 3, pp. 223-233)


Chen Shao, Feng Gao, Xiaozhong Hu, Khaled A. Al-Rasheid and Allan Warren
Ontogenesis and Molecular Phylogeny of a New Marine Urostylid Ciliate, Anteholosticha petzi n. sp. (Ciliophora, Urostylida)” (vol. 58, Issue 3, pp. 254-265)

Regula Schmid-Hempel and Martina Tognazzo
Molecular Divergence Defines Two Distinct Lineages of Chritidia bombi (Trypanosomatidae), Parasites of Bumblebees” (vol. 57, Issue 4, pp. 337-345)

Y. P. Chen, J. D. Evans, C. Murphy, R. Gutell, M. Zucker, D. Gundensen-Rindal & J. S. Pettis
Morphological, Molecular and Phylogenetic Characterization of Nosema ceranae, a Microsporidian Parasite Isolated from the European Honey Bee, Apis mellifera” (vol. 56, Issue 2, pp. 142-147)

S. Kim, M. G. Park, K.-Y. Kim, C.-H. Kim, W. Yih, J. S. Park & D. W. Coats
Genetic Diversity of Parasitic Dinoflagellates in the Genus Amoebophyra and Its Relationship to Parasite Biology and Biogeography” (vol. 55, Issue 1, pp. 1-8)

S.L. Schmidt, D. Bernhard, M. Schlegel & W. Foissner.
Phylogeny of the Stichotrichia (Ciliophora: Spirotrichea) Reconstructed with Nuclear Small Subunit rRNA Gene Sequences: Discrepancies and Accordances with Morphological Data” (vol. 54, Issue 2, pp. 201-209)

D. Barth, S. Krenek, S. I. Fokin & T. U. Berendonk.
Intraspecific Genetic Variation in Paramecium Revealed by Mitochondrial Cytochrome c Oxidase I Sequences” (vol. 53, Issue 1, pp. 20-25)

C. D. Lowe, A. Day, S. J. Kemp & D. J. S. Montagnes.
There are High Levels of Functional and Genetic Diversity in Oxyrrhis marina” (vol. 52, Issue 3, pp. 250-257)

C. H. Slamovits, B. A. P. Williams, & P. J. Keeling.
Transfer of Nosema locustae (Microsporidia) to Antonospora locustae n. comb. Based on Molecular and Ultrastructural Data” (vol. 51, Issue 2, pp. 207-213)

Photo by David J. Patterson

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